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Supported by grants from the Beijing Science and Technology Planning Project (CN) (D17110002617002), National Key Research and Development Program of China (2016YFC1302801), National Key Research and Development Program of China (2018YFC1313103), CAMS Innovation Fund for Medical Sciences (2016-I2M-1-001), (2017-I2M-1-106), and (2019-I2M-2-004), Sanming Project of Medicine in Shenzhen (No. SZSM201911008), and PUMC Youth Fund and the Fundamental Research Funds for the Central Universities (2017320012).